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AcknowledgementsIt would have been impossible to complete this PhD entirely on my own. I have collaborated with and received support from numerous people. While I cannot list everyone by name, I want to express my gratitude for your support. I want to give special thanks to the following people:Mariska Kret, thank you for the opportunities and encouragement you provided, allowing me to develop and pursue my main interests and skills throughout my PhD. The lessons and skills I’ve acquired under your supervision will stay with me forever. Yena Kim, your support during the first orang-utan touchscreen study was crucial, and gave me the confidence to continue. Juan Olvido Perea-García, even though you were not initially involved in my project, there are few people who have read so much of the content of my thesis. Your feedback was invaluable and I really enjoyed our collaborative research projects. My paranymphs Iliana Samara and Julia Folz; Iliana, we have collaborated on many projects over the years, and half of this thesis would not even exist without your contributions. I particularly cherished our Skype-statistics-writing sessions during the first COVID lockdown. They really brightened my days during a difficult and insecure period. Julia, we started our PhDs around the same time and I found it really valuable to share experiences and occasional frustrations over a nice beer. Evy van Berlo, my studies have been built upon the solid foundation you laid during your PhD. I really enjoyed collaborating and our summerschoolroadtrip in Germany is one of the fondest memories of my PhD. Tonko Zijlstra, thank you for your humour and calm advice when I needed it. I would also like to extend my gratitude to all the members of the CoPAN lab for making my PhD experience so enjoyable. Furthermore, I want to thank SOLO Research and Lab Support, especially Evert Dekker and Iris Spruit, for their technical support throughout my research projects.This project would have been impossible without Thomas Bionda. Thomas, thank you for your support and your trust in me. You always made me feel welcome in Apenheul, and gave me the confidence that I was the right person for this project, even when I was doubting myself. Lisette van den Berg, thank you for your efforts to enable me and many other researchers to perform research in Apenheul. José Kok, thank you for giving us the opportunity to perform orangutan research in Ouwehands Dierenpark. Furthermore, I want to thank all of the caretakers who made it possible to do research with the orang-utans. In Apenheul: Acknowledgements5Tom Roth.indd 5 08-01-2024 10:41